拟阿勇蛞蝓科

柄眼目的一科软体动物

拟阿勇蛞蝓科(学名:Ariophantidae[2],又名丝鳖甲蜗牛科[3],在生物分类学是会呼吸空气的陆地蜗牛英语land snail蛞蝓的一个陆生英语terrestrial molluscs肺螺类腹足纲软体动物支序的成员,属于柄眼目鳖甲蜗牛总科英语Helicarionoidea(Helicarionoidea)[4][5]

拟阿勇蛞蝓科
两个活体的印度巨楯蛞蝓英语Macrochlamys indica
科学分类 编辑
界: 动物界 Animalia
门: 软体动物门 Mollusca
纲: 腹足纲 Gastropoda
目: 柄眼目 Stylommatophora
亚目: 旋蜗牛亚目 Helicina
下目: 蛞蝓下目 Limacoidei
总科: 勇蜗总科 Helicarionoidea
科: 拟阿勇蛞蝓科 Ariophantidae
Godwin-Austen英语Henry Haversham Godwin-Austen, 1888[1]

见内文

分布

根据Hausdorf (2000),本科物种只分布于印度英语List of non-marine molluscs of India和东南亚[6];但其实本科物种大多是原产于非洲、东南亚和新几内亚。

系统分类学

亚科

根据2005年《布歇特和洛克罗伊的腹足类分类》,本科由下列三个亚科组成[4]

  • Ariophantinae Godwin-Austen, 1888 - synonyms: Naninidae Pfeffer, 1878 (inv.); Hemiplectinae Gude & B. B. Woodward, 1921[7]
  • Macrochlamydinae Godwin-Austen, 1888[1] - synonyms: Tanychlamydinae H. B. Baker, 1928;[8] Vitrinulini Schileyko, 2003[9]
  • Ostracolethinae Simroth英语Heinrich Simroth, 1901[10] - synonyms: Myotestidae Collinge, 1902;[11] Parmarioninae Godwin-Austen, 1908;[12] Laocaiini Schileyko, 2002;[13] Microparmarionini Schileyko, 2003[14]

截至2020年2月29日 (2020-02-29)MolluscaBase纪录本科包括下列各亚科与属:

  • 真奥氏蛞蝓属 Euaustenia T. D. A. Cockerell, 1891
  • 巨楯蛞蝓属 Macrochlamys Gray, 1847[18]:本亚科的模式属[4],又名巨丝鳖甲蜗牛属[3]
    • Macrochlamys (Bensonies) aborensis (Godwin-Austen, 1918)
    • Macrochlamys albulus Godwin-Austen, 1918
    • Macrochlamys alluaudi Bavay & Dautzenberg, 1900
    • Macrochlamys amboinensis (E.von Martens, 1864)
    • Macrochlamys amdoana Möllendorff, 1899
    • Macrochlamys anceyi Degner, 1928
    • Macrochlamys andersoniana (G.Nevill, 1877)
    • Macrochlamys angigyra Yen, 1939
    • Macrochlamys angigyrata Godwin-Austen, 1910
    • Macrochlamys angulatus Möllendorff, 1887
    • Macrochlamys anonae Godwin-Austen, 1883
    • Macrochlamys apex Möllendorff, 1888
    • Macrochlamys aspides (W.H.Benson, 1863)
    • Macrochlamys atoma W.T.Blanford, 1904
    • Macrochlamys atricolor (Godwin-Austen, 1875)
    • Macrochlamys aufschnaiteri Blume, 1957
    • Macrochlamys aulopis (W.H.Benson, 1863)
    • Macrochlamys aurantia Pholyotha & Panha, 2018
    • Macrochlamys bapuensis Godwin-Austen, 1918
    • Macrochlamys bartoni de Morgan, 1885
    • Macrochlamys battlimalvensis Godwin-Austen, 1888
    • Macrochlamys beata Godwin-Austen, 1907
    • Macrochlamys bilineata (Godwin-Austen, 1876)
    • Macrochlamys boettgeri (Hilber, 1883)
    • Macrochlamys brachystia Pholyotha, Sutcharit, Tongkerd, Lin & Panha, 2020
    • Macrochlamys brunnea Möllendorff, 1902
    • Macrochlamys burkilii Godwin-Austen, 1918
    • Macrochlamys buruana Möllendorff, 1902
    • Macrochlamys cacharica Godwin-Austen, 1883
    • Macrochlamys cagayanica Quadras & Möllendorff, 1893
    • Macrochlamys callojuncta Ancey, 1898
    • Macrochlamys castaneolabiata Godwin-Austen, 1883
    • Macrochlamys cathaiana Möllendorff, 1899
    • Macrochlamys causia (W.H.Benson, 1859)
    • Macrochlamys caverna Pholyotha & Panha, 2018
    • Macrochlamys ceromatica (Morelet, 1881)
    • Macrochlamys chaos W.T.Blanford, 1904
    • Macrochlamys choinix (W.H.Benson, 1861)
    • Macrochlamys cincta Möllendorff, 1882
    • Macrochlamys cinctula Godwin-Austen, 1910
    • Macrochlamys clessini Westerlund, 1902
    • Macrochlamys coeligana Collinge, 1902
    • Macrochlamys coleus Pholyotha & Panha, 2018
    • Macrochlamys corusca (Morelet, 1877)
    • Macrochlamys (Macrochlamys) crebristriata C.Semper, 1870
    • Macrochlamys curvilabris W.T.Blanford, 1904
    • Macrochlamys dalingensis Godwin-Austen, 1883
    • Macrochlamys dammeriana Möllendorff, 1899
    • Macrochlamys damsangensis Godwin-Austen, 1907
    • Macrochlamys darjilingensis Godwin-Austen, 1883
    • Macrochlamys davidi (Deshayes, 1874)
    • Macrochlamys declivis Möllendorff, 1901
    • Macrochlamys decursus Godwin-Austen, 1914
    • Macrochlamys decussata (W.H.Benson, 1836)
    • Macrochlamys dikrangensis Godwin-Austen, 1910
    • Macrochlamys discus Möllendorff, 1884
    • Macrochlamys dohertyi E.A.Smith, 1897
    • Macrochlamys dolani Pilsbry, 1934
    • Macrochlamys dorani Godwin-Austen, 1883
    • Macrochlamys douvillei Dautzenberg & H.Fischer, 1906
    • Macrochlamys enata Godwin-Austen, 1883
    • Macrochlamys euryomphala (Möllendorff, 1898)
    • Macrochlamys euspira Möllendorff, 1901
    • Macrochlamys euterpe (L.Pfeiffer, 1861)
    • Macrochlamys evidens Godwin-Austen, 1910
    • Macrochlamys excepta (Mabille, 1887)
    • Macrochlamys extraria Godwin-Austen, 1910
    • Macrochlamys exul (Theobald, 1864)
    • Macrochlamys faceta Godwin-Austen, 1883
    • Macrochlamys fargesiana (Heude, 1885)
    • Macrochlamys (Parvatella) flemingi (L.Pfeiffer, 1857)
    • Macrochlamys fordiana Godwin-Austen, 1888
    • Macrochlamys fragilis (T.Hutton, 1838)
    • Macrochlamys fragosus Godwin-Austen, 1907
    • Macrochlamys fulvus H.Rolle, 1908
    • Macrochlamys ganjamensis Godwin-Austen, 1910
    • Macrochlamys garoensis Godwin-Austen, 1910
    • Macrochlamys geoffreyi (H.Adams, 1868)
    • Macrochlamys glauca (L.Pfeiffer, 1846)
    • Macrochlamys glyptorhaphe Möllendorff, 1901
    • Macrochlamys godwini Tryon, 1886
    • Macrochlamys gurhwalensis (Godwin-Austen, 1899)
    • Macrochlamys hardwickei Godwin-Austen, 1883
    • Macrochlamys hatchongi de Morgan, 1885
    • Macrochlamys hebescens (W.T.Blanford, 1866)
    • Macrochlamys hendersoni Ancey, 1905
    • Macrochlamys hengdanensis Godwin-Austen, 1907
    • Macrochlamys henrici C.Semper, 1870
    • Macrochlamys hepatizon Godwin-Austen, 1888
    • Macrochlamys herbia Godwin-Austen, 1910
    • Macrochlamys hippocastaneum Godwin-Austen, 1918
    • Macrochlamys hodgsoni (W.H.Benson, 1859)
    • Macrochlamys hookeri Godwin-Austen, 1907
    • Macrochlamys hunancola (Möllendorff, 1887)
    • Macrochlamys igorrotica (Möllendorff, 1896)
    • Macrochlamys imperforata Schepman, 1918
    • Macrochlamys indica Godwin-Austen, 1883
    • Macrochlamys indifferens E.A.Smith, 1896
    • Macrochlamys infans (Reeve, 1854)
    • Macrochlamys infausta (W.T.Blanford, 1866)
    • Macrochlamys jainiana Godwin-Austen, 1883
    • Macrochlamys japvoensis Godwin-Austen, 1910
    • Macrochlamys kandiensis Godwin-Austen, 1883
    • Macrochlamys kashmirensis (G.Nevill, 1878)
    • Macrochlamys kasnakowi (Westerlund, 1898)
    • Macrochlamys kelantanensis Möllendorff, 1902
    • Macrochlamys kempi Godwin-Austen, 1883
    • Macrochlamys koliaensis Godwin-Austen, 1883
    • Macrochlamys kuluensis W.T.Blanford, 1904
    • Macrochlamys kurtzi Godwin-Austen, 1910
    • Macrochlamys lahupaensis Godwin-Austen, 1907
    • Macrochlamys latus Godwin-Austen, 1888
    • Macrochlamys lecythis (W.H.Benson, 1852)
    • Macrochlamys leggeae Sajan, Tripathy, Chandra & Sivakumar, 2019
    • Macrochlamys lemma Pholyotha & Panha, 2018
    • Macrochlamys lemna Pholyotha & Panha, 2018
    • Macrochlamys lhotaensis Godwin-Austen, 1883
    • Macrochlamys lixa (W.T.Blanford, 1866)
    • Macrochlamys loana (Gredler, 1882)
    • Macrochlamys longicauda Godwin-Austen, 1883
    • Macrochlamys lubrica (W.H.Benson, 1852)
    • Macrochlamys lubricata Godwin-Austen, 1910
    • Macrochlamys lumsdeni (Godwin-Austen, 1910)
    • Macrochlamys luyorensis Godwin-Austen, 1918
    • Macrochlamys luzonica (Möllendorff, 1894)
    • Macrochlamys mahadeoensis Godwin-Austen, 1907
    • Macrochlamys malaccana (L.Pfeiffer, 1855)
    • Macrochlamys martini Schepman, 1893
    • Macrochlamys masoni Godwin-Austen, 1910
    • Macrochlamys mentaweiense Degner, 1928
    • Macrochlamys minuta E.von Martens, 1891
    • Macrochlamys molecula (W.H.Benson, 1859)
    • Macrochlamys moupiniana (Deshayes, 1870)
    • Macrochlamys munipurensis Godwin-Austen, 1907
    • Macrochlamys murdochi Godwin-Austen, 1918
    • Macrochlamys neherensis (W.H.Benson, 1864)
    • Macrochlamys nemothaensis Godwin-Austen, 1910
    • Macrochlamys nengloensis Godwin-Austen, 1883
    • Macrochlamys nepos (L.Pfeiffer, 1855)
    • Macrochlamys nitidissima Möllendorff, 1883
    • Macrochlamys notha W.T.Blanford, 1904
    • Macrochlamys novoguineensis Schepman, 1919
    • Macrochlamys noxia W.T.Blanford, 1904
    • Macrochlamys nuda (Reeve, 1852)
    • Macrochlamys obiana Möllendorff, 1902
    • Macrochlamys opiparus Godwin-Austen, 1883
    • Macrochlamys originaria Godwin-Austen, 1883
    • Macrochlamys pacata Godwin-Austen, 1883
    • Macrochlamys papuana E.A.Smith, 1896
    • Macrochlamys papuensis Preston, 1902
    • Macrochlamys par Schmacker & Boettger, 1891
    • Macrochlamys patane (W.H.Benson, 1859)
    • Macrochlamys patens W.T.Blanford, 1904
    • Macrochlamys paurhiensis (Godwin-Austen, 1899)
    • Macrochlamys pedina (W.H.Benson, 1865)
    • Macrochlamys pedisequae Heude, 1886
    • Macrochlamys perfragilis Godwin-Austen, 1907
    • Macrochlamys perfucata (W.H.Benson, 1853)
    • Macrochlamys perincospicua Godwin-Austen, 1888
    • Macrochlamys peringundensis R.H.Beddome, 1891
    • Macrochlamys perplana Godwin-Austen, 1883
    • Macrochlamys perrotteti (L.Pfeiffer, 1851)
    • Macrochlamys petasus (W.H.Benson, 1859)
    • Macrochlamys petrosa (T.Hutton, 1834)
    • Macrochlamys planata (Heude, 1882)
    • Macrochlamys planiuscula (W.H.Benson, 1838)
    • Macrochlamys planorbiformis Bollinger, 1918
    • Macrochlamys planula (Heude, 1882)
    • Macrochlamys plicifera (W.T.Blanford, 1881)
    • Macrochlamys politula Godwin-Austen, 1883
    • Macrochlamys prava W.T.Blanford, 1905
    • Macrochlamys proba Godwin-Austen, 1910
    • Macrochlamys propinqua Godwin-Austen, 1910
    • Macrochlamys pseudaulopis Godwin-Austen, 1888
    • Macrochlamys pseudochoinix W.T.Blanford, 1904
    • Macrochlamys pseudosuccinea E.A.Smith, 1894
    • Macrochlamys pseustes E.A.Smith, 1893
    • Macrochlamys psittacinus Godwin-Austen, 1918
    • Macrochlamys psyche Vermeulen, Luu, Theary & Anker, 2019
    • Macrochlamys quadrasi (Möllendorff, 1896)
    • Macrochlamys quesita Godwin-Austen, 1910
    • Macrochlamys rabani Godwin-Austen, 1910
    • Macrochlamys radia Godwin-Austen, 1910
    • Macrochlamys rakaensis Godwin-Austen, 1907
    • Macrochlamys ramburianus (Mabille & Le Mesle, 1866)
    • Macrochlamys razamiensis Godwin-Austen, 1907
    • Macrochlamys rejecta (L.Pfeiffer, 1859)
    • Macrochlamys rejectella (Mabille, 1889)
    • Macrochlamys renitens (Morelet, 1861)
    • Macrochlamys resinacea (Heude, 1885)
    • Macrochlamys richilaensis Godwin-Austen, 1907
    • Macrochlamys roberti Godwin-Austen, 1883
    • Macrochlamys robsoni Rensch, 1932
    • Macrochlamys rorida (W.H.Benson, 1859)
    • Macrochlamys rotungensis Godwin-Austen, 1918
    • Macrochlamys rubellocincta (W.T.Blanford, 1870)
    • Macrochlamys russeola (Morelet, 1851)
    • Macrochlamys rusticula Godwin-Austen, 1883
    • Macrochlamys rutila W.T.Blanford, 1905
    • Macrochlamys sacrata Godwin-Austen, 1910
    • Macrochlamys salmonea (Ancey, 1882)
    • Macrochlamys saltus Godwin-Austen, 1910
    • Macrochlamys salwinensis Godwin-Austen, 1907
    • Macrochlamys sanchezi Quadras & Möllendorff, 1894
    • Macrochlamys sanctijohni (Godwin-Austen, 1891)
    • Macrochlamys sata Godwin-Austen, 1883
    • Macrochlamys sathilaensis Godwin-Austen, 1907
    • Macrochlamys scyphus Godwin-Austen, 1910
    • Macrochlamys segnis Pilsbry, 1934
    • Macrochlamys sequax (W.H.Benson, 1859)
    • Macrochlamys sequius Godwin-Austen, 1907
    • Macrochlamys shengorensis Godwin-Austen, 1883
    • Macrochlamys shimangensis Godwin-Austen, 1916
    • Macrochlamys shisha (Godwin-Austen, 1875)
    • Macrochlamys silcuriensis Godwin-Austen, 1914
    • Macrochlamys silvatica Godwin-Austen, 1910
    • Macrochlamys simplex Heude, 1890
    • Macrochlamys sinensis (Heude, 1882)
    • Macrochlamys sinica E.von Martens, 1875
    • Macrochlamys sphaerica Godwin-Austen, 1910
    • Macrochlamys spiralifer Vermeulen, 1996
    • Macrochlamys spiriplana (Gredler, 1882)
    • Macrochlamys spreta W.T.Blanford, 1904
    • Macrochlamys stenogyra Möllendorff, 1901
    • Macrochlamys stephoides Stoliczka, 1873
    • Macrochlamys stephus (W.H.Benson, 1861)
    • Macrochlamys strephoides Stoliczka, 1873
    • Macrochlamys striata Godwin-Austen, 1910
    • Macrochlamys striaticostata Godwin-Austen, 1910
    • Macrochlamys stumpfii O.Boettger, 1889
    • Macrochlamys subangulata Godwin-Austen, 1910
    • Macrochlamys subcornea (L.Pfeiffer, 1861)
    • Macrochlamys subcorpulenta Vermeulen & Liew, 2022
    • Macrochlamys subjecta (W.H.Benson, 1852)
    • Macrochlamys subpetasus Godwin-Austen, 1888
    • Macrochlamys sufflava Godwin-Austen, 1910
    • Macrochlamys superflua W.T.Blanford, 1904
    • Macrochlamys superlita (Morelet, 1862)
    • Macrochlamys sylhetensis Godwin-Austen, 1910
    • Macrochlamys tanirensis Godwin-Austen, 1883
    • Macrochlamys tanymentula Pholyotha & Panha, 2018
    • Macrochlamys tecta (Souleyet, 1852)
    • Macrochlamys tenuiarata Vermeulen & Liew, 2022
    • Macrochlamys tenuicula H.Adams, 1868
    • Macrochlamys tenuigranosa Dautzenberg, 1894
    • Macrochlamys terminalis Godwin-Austen, 1910
    • Macrochlamys terminus Godwin-Austen, 1899
    • Macrochlamys tersa (Issel, 1874)
    • Macrochlamys thaitieni Thach, 2021
    • Macrochlamys theobaldi (Godwin-Austen, 1888)
    • Macrochlamys tratanensis (Jousseaume, 1894)
    • Macrochlamys trilobata Vermeulen & Liew, 2022
    • Macrochlamys trusmadiensis Vermeulen & Liew, 2022
    • Macrochlamys tugurium (W.H.Benson, 1852)
    • Macrochlamys turanica E.von Martens, 1874
    • Macrochlamys udus Godwin-Austen, 1899
    • Macrochlamys umbraticola Godwin-Austen, 1883
    • Macrochlamys umbrina (Reeve, 1854)
    • Macrochlamys vallicola (Reeve, 1854)
    • Macrochlamys vertex Godwin-Austen, 1910
    • Macrochlamys vesica Godwin-Austen, 1910
    • Macrochlamys vesicula (W.H.Benson, 1838)
    • Macrochlamys vitrinoides (Deshayes, 1831)
    • Macrochlamys woodiana (Reeve, 1852)
    • Macrochlamys woodmasoni Godwin-Austen, 1888
    • Macrochlamys zemoensis Godwin-Austen, 1907
    • Macrochlamys zero (Mabille, 1887)
  • 拟小囊螺属 Microcystina Mörch, 1872
  • Oxytesta Zilch, 1956
  • 罗氏螺属 Rahula Godwin-Austen, 1907
  • 萨氏螺属 Sarika Godwin-Austen, 1907
  • 赛氏螺属 Sesara Albers, 1860
  • Syama W. T. Blanford & Godwin-Austen, 1908[19]
  • 巨楯蛞蝓属 Tanychlamys Benson, 1834[4]
  • Vitrinula Gray, 1857[4][20]
 
Microparmarion simrothi

小黄壳蛞蝓亚科 Ostracolethinae Simroth, 1901

  • Cryptosemelus Collinge, 1902
    • Cryptosemelus betarmon Pholyotha, 2021[21]
    • Cryptosemelus gracilis Collinge, 1902
    • Cryptosemelus niger S.Tumpeesuwan & C.Tumpeesuwan, 2023
    • Cryptosemelus tigrinus Pholyotha, 2021[21]
  • Laocaia Kuzminykh, 1999[4]
  • Microparmarion Simroth, 1893[4]
    • Microparmarion andamanica Collinge, 1901
    • Microparmarion austeni Simroth, 1893
    • Microparmarion basifixus Vermeulen & Liew, 2022
    • Microparmarion boettgeri Simroth, 1898
    • Microparmarion bruneopallescens Collinge, 1901
    • Microparmarion convolutus Vermeulen & Liew, 2022
    • Microparmarion exquadratus Schilthuizen, Clavera, Khoo, Bondar, Elder, Bouma, Eddington, Reisinger, Cosentino, Rossato & Delledonne, 2019
    • Microparmarion fruhstorferi Simroth, 1898
    • Microparmarion litteratus Schepman, 1896
    • Microparmarion malayanus (Collinge, 1903)
    • Microparmarion moellendorffi Collinge, 1901
    • Microparmarion pollonerai Collinge & Godwin-Austen, 1895
    • Microparmarion sallehi H.Wu, Ezzwan & Hamdani, 2023[22]
    • Microparmarion simrothi Collinge & Godwin-Austen, 1895
    • Microparmarion strubelli Simroth, 1893
  • 米农螺属 Minyongia Godwin-Austen, 1916
    • Minyongia kempi Godwin-Austen, 1916
  • Myotesta Collinge, 1901[4]
  • Ostracolethe Simroth, 1901 - 为Ostracolethinae亚科的模式属 [4]
  • 鳖甲蛞蝓属 Parmarion P. Fischer, 1855,盾勇螺属[3][4]
  • 菲律宾蛞蝓属 Philippinella Möllendorff, 1899
    • Philippinella carinata (Möllendorff, 1894)
    • Philippinella moellendorffi Collinge, 1899
    • Philippinella quadrasi (Möllendorff, 1894)

亲缘支序图

以下的系统发生学分支图显示了根据Hausdorf (2000)所描述蛞蝓下目各科之间的亲缘关系[6]

 蛞蝓下目 Limacoidei 
 Staffordioidea英语Staffordioidea 

Staffordiidae

 Dyakioidea英语Dyakioidea 

Dyakiidae

Pristilomatidae英语Pristilomatidae

克氏螺科 Chronidae

真蛞蝓科 Euconulidae

Trochomorphidae英语Trochomorphidae

腹齿螺科 Gastrodontidae

Oxychilidae英语Oxychilidae

 Parmacelloidea英语Parmacelloidea 

Trigonochlamydidae英语Trigonochlamydidae

Parmacellidae英语Parmacellidae

Milacidae英语Milacidae

 带螺总科 Zonitoidea 

带螺科 Zonitidae

鳖甲蜗牛科 Helicarionidae

拟阿勇蛞蝓科 Ariophantidae

Urocyclidae英语Urocyclidae

 Limacoidea英语Limacoidea 

Vitrinidae英语Vitrinidae

Boettgerillidae英语Boettgerillidae

蛞蝓科 Limacidae

野蛞蝓科 Agriolimacidae

参考文献

  1. ^ 1.0 1.1 Godwin-Austen H. H. (April 1888). Land and freshwater Mollusca of India. including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean, supplementary to Messrs. Theobald and Hanley's Conchologia Indica. Taylor & Francis, London, volume 1, part 6: 207-257, page 253.
  2. ^ 中国科学院动物研究所. 真锥恰里螺. 《中国动物物种编目数据库》. 中国科学院微生物研究所. [2009-04-28]. (原始内容存档于2016-03-05). 
  3. ^ 3.0 3.1 3.2 Ariophantidae 絲鼈甲蝸牛科. 台湾生物多样性资讯入口网. [2020-03-02]. (原始内容存档于2020-08-13) (中文(繁体)). 
  4. ^ 4.00 4.01 4.02 4.03 4.04 4.05 4.06 4.07 4.08 4.09 4.10 Bouchet, P.; Rocroi, J.-P.; Frýda, J.; Hausdorf, B.; Ponder, W.; Valdés, Á. & Warén, A. Classification and nomenclator of gastropod families. Malacologia (Hackenheim, Germany: ConchBooks). 2005, 47 (1–2): 1–397. ISBN 3-925919-72-4. OCLC 61351079. OL 42989175M. S2CID 83847814. 
  5. ^ Bouchet, P.; Rocroi, J.-P.; Hausdorf, B.; Kaim, A.; Kano, Y.; Nützel, A.; Parkhaev, P.; Schrödl, M.; Strong, E.E. Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families. Malacologia. 2017, 61 (1–2): 1–526. OCLC 7296100651. S2CID 91051256. doi:10.4002/040.061.0201. 
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延伸阅读

  • Schileyko A. A. (2003). "Treatise on recent terrestrial pulmonate mollusks. 10. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae". Ruthenica, Supplement 2. 1309-1466.

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