拟阿勇蛞蝓科
柄眼目的一科软体动物
（重定向自Ariophantidae）
拟阿勇蛞蝓科（學名：Ariophantidae）[2]，又名絲鼈甲蝸牛科[3]，在生物分类学是會呼吸空氣的陸地蝸牛及蛞蝓的一個陸生肺螺類腹足纲软体动物支序的成員，屬於柄眼目鳖甲蜗牛總科（Helicarionoidea）[4][5]。
拟阿勇蛞蝓科
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| 两個活体的印度巨楯蛞蝓 | |
科学分类 
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| 界： | 动物界 Animalia |
| 门： | 軟體動物門 Mollusca |
| 纲： | 腹足纲 Gastropoda |
| 目： | 柄眼目 Stylommatophora |
| 亚目： | 旋蝸牛亞目 Helicina |
| 下目： | 蛞蝓下目 Limacoidei |
| 总科： | 勇蜗总科 Helicarionoidea |
| 科： | 拟阿勇蛞蝓科 Ariophantidae Godwin-Austen, 1888[1] |
| 屬 | |
分佈
根據Hausdorf (2000)，本科物種只分佈於印度和東南亞[6]；但其實本科物種大多是原產於非洲、東南亞和新幾內亞。
系統分類學
亞科
根據2005年《布歇特和洛克羅伊的腹足類分類》，本科由下列三個亞科組成[4]：
- Ariophantinae Godwin-Austen, 1888 - synonyms: Naninidae Pfeffer, 1878 (inv.); Hemiplectinae Gude & B. B. Woodward, 1921[7]
- Macrochlamydinae Godwin-Austen, 1888[1] - synonyms: Tanychlamydinae H. B. Baker, 1928;[8] Vitrinulini Schileyko, 2003[9]
- Ostracolethinae Simroth, 1901[10] - synonyms: Myotestidae Collinge, 1902;[11] Parmarioninae Godwin-Austen, 1908;[12] Laocaiini Schileyko, 2002;[13] Microparmarionini Schileyko, 2003[14]
屬
截至2020年2月29日[update]，MolluscaBase紀錄本科包括下列各亞科與屬：
拟阿勇蛞蝓亚科 Ariophantinae Godwin-Austen, 1883
- 拟阿勇蛞蝓属 Ariophanta Des Moulins, 1829：也是本科的模式屬[4]
- Bapuia Godwin-Austen, 1918
- 本氏蛞蝓属 Bensonies H. B. Baker, 1938
- Cryptogirasia Cockerell, 1891
- 隐带螺属 Cryptozona Mörch, 1872
- Dalingia Godwin-Austen, 1907
- Dekhania Godwin-Austen, 1887
- Dihangia Godwin-Austen, 1916
- 真卷螺属 Euplecta C. Semper, 1870
- Galongia Godwin-Austen, 1916
- 半卷螺属 Hemiplecta Albers, 1850
- 因陀罗蜗牛属 Indrella Godwin-Austen, 1901: 其学名来自于印度教之神因陀罗[15]:190
- 卡西山螺属 Khasiella Godwin-Austen, 1899
- Koratia Godwin-Austen, 1919
- 玛蛞蝓属 Mariaella Gray, 1855，见于台湾
- 巨奥氏蛞蝓属 Megaustenia Cockerell, 1912
- Ratnadvipia Godwin-Austen, 1899[16]：斯里蘭卡原生種[17]
- Ravana Godwin-Austen, 1901
- Schwammeria Schileyko, 2010
- Sitalinopsis Thiele, 1931
- Taphrospira W.T. Blanford, 1905
- Xesta Albers, 1850
- 光螺属 Xestina Pfeffer, 1878，为隐带螺属（Cryptozona Mörch, 1872 ）的异名
- 真奥氏蛞蝓属 Euaustenia T. D. A. Cockerell, 1891
- 巨楯蛞蝓属 Macrochlamys Gray, 1847[18]：本亞科的模式屬[4]，又名巨絲鼈甲蝸牛屬[3]。
- Macrochlamys (Bensonies) aborensis (Godwin-Austen, 1918)
- Macrochlamys albulus Godwin-Austen, 1918
- Macrochlamys alluaudi Bavay & Dautzenberg, 1900
- Macrochlamys amboinensis (E.von Martens, 1864)
- Macrochlamys amdoana Möllendorff, 1899
- Macrochlamys anceyi Degner, 1928
- Macrochlamys andersoniana (G.Nevill, 1877)
- Macrochlamys angigyra Yen, 1939
- Macrochlamys angigyrata Godwin-Austen, 1910
- Macrochlamys angulatus Möllendorff, 1887
- Macrochlamys anonae Godwin-Austen, 1883
- Macrochlamys apex Möllendorff, 1888
- Macrochlamys aspides (W.H.Benson, 1863)
- Macrochlamys atoma W.T.Blanford, 1904
- Macrochlamys atricolor (Godwin-Austen, 1875)
- Macrochlamys aufschnaiteri Blume, 1957
- Macrochlamys aulopis (W.H.Benson, 1863)
- Macrochlamys aurantia Pholyotha & Panha, 2018
- Macrochlamys bapuensis Godwin-Austen, 1918
- Macrochlamys bartoni de Morgan, 1885
- Macrochlamys battlimalvensis Godwin-Austen, 1888
- Macrochlamys beata Godwin-Austen, 1907
- Macrochlamys bilineata (Godwin-Austen, 1876)
- Macrochlamys boettgeri (Hilber, 1883)
- Macrochlamys brachystia Pholyotha, Sutcharit, Tongkerd, Lin & Panha, 2020
- Macrochlamys brunnea Möllendorff, 1902
- Macrochlamys burkilii Godwin-Austen, 1918
- Macrochlamys buruana Möllendorff, 1902
- Macrochlamys cacharica Godwin-Austen, 1883
- Macrochlamys cagayanica Quadras & Möllendorff, 1893
- Macrochlamys callojuncta Ancey, 1898
- Macrochlamys castaneolabiata Godwin-Austen, 1883
- Macrochlamys cathaiana Möllendorff, 1899
- Macrochlamys causia (W.H.Benson, 1859)
- Macrochlamys caverna Pholyotha & Panha, 2018
- Macrochlamys ceromatica (Morelet, 1881)
- Macrochlamys chaos W.T.Blanford, 1904
- Macrochlamys choinix (W.H.Benson, 1861)
- Macrochlamys cincta Möllendorff, 1882
- Macrochlamys cinctula Godwin-Austen, 1910
- Macrochlamys clessini Westerlund, 1902
- Macrochlamys coeligana Collinge, 1902
- Macrochlamys coleus Pholyotha & Panha, 2018
- Macrochlamys corusca (Morelet, 1877)
- Macrochlamys (Macrochlamys) crebristriata C.Semper, 1870
- Macrochlamys curvilabris W.T.Blanford, 1904
- Macrochlamys dalingensis Godwin-Austen, 1883
- Macrochlamys dammeriana Möllendorff, 1899
- Macrochlamys damsangensis Godwin-Austen, 1907
- Macrochlamys darjilingensis Godwin-Austen, 1883
- Macrochlamys davidi (Deshayes, 1874)
- Macrochlamys declivis Möllendorff, 1901
- Macrochlamys decursus Godwin-Austen, 1914
- Macrochlamys decussata (W.H.Benson, 1836)
- Macrochlamys dikrangensis Godwin-Austen, 1910
- Macrochlamys discus Möllendorff, 1884
- Macrochlamys dohertyi E.A.Smith, 1897
- Macrochlamys dolani Pilsbry, 1934
- Macrochlamys dorani Godwin-Austen, 1883
- Macrochlamys douvillei Dautzenberg & H.Fischer, 1906
- Macrochlamys enata Godwin-Austen, 1883
- Macrochlamys euryomphala (Möllendorff, 1898)
- Macrochlamys euspira Möllendorff, 1901
- Macrochlamys euterpe (L.Pfeiffer, 1861)
- Macrochlamys evidens Godwin-Austen, 1910
- Macrochlamys excepta (Mabille, 1887)
- Macrochlamys extraria Godwin-Austen, 1910
- Macrochlamys exul (Theobald, 1864)
- Macrochlamys faceta Godwin-Austen, 1883
- Macrochlamys fargesiana (Heude, 1885)
- Macrochlamys (Parvatella) flemingi (L.Pfeiffer, 1857)
- Macrochlamys fordiana Godwin-Austen, 1888
- Macrochlamys fragilis (T.Hutton, 1838)
- Macrochlamys fragosus Godwin-Austen, 1907
- Macrochlamys fulvus H.Rolle, 1908
- Macrochlamys ganjamensis Godwin-Austen, 1910
- Macrochlamys garoensis Godwin-Austen, 1910
- Macrochlamys geoffreyi (H.Adams, 1868)
- Macrochlamys glauca (L.Pfeiffer, 1846)
- Macrochlamys glyptorhaphe Möllendorff, 1901
- Macrochlamys godwini Tryon, 1886
- Macrochlamys gurhwalensis (Godwin-Austen, 1899)
- Macrochlamys hardwickei Godwin-Austen, 1883
- Macrochlamys hatchongi de Morgan, 1885
- Macrochlamys hebescens (W.T.Blanford, 1866)
- Macrochlamys hendersoni Ancey, 1905
- Macrochlamys hengdanensis Godwin-Austen, 1907
- Macrochlamys henrici C.Semper, 1870
- Macrochlamys hepatizon Godwin-Austen, 1888
- Macrochlamys herbia Godwin-Austen, 1910
- Macrochlamys hippocastaneum Godwin-Austen, 1918
- Macrochlamys hodgsoni (W.H.Benson, 1859)
- Macrochlamys hookeri Godwin-Austen, 1907
- Macrochlamys hunancola (Möllendorff, 1887)
- Macrochlamys igorrotica (Möllendorff, 1896)
- Macrochlamys imperforata Schepman, 1918
- Macrochlamys indica Godwin-Austen, 1883
- Macrochlamys indifferens E.A.Smith, 1896
- Macrochlamys infans (Reeve, 1854)
- Macrochlamys infausta (W.T.Blanford, 1866)
- Macrochlamys jainiana Godwin-Austen, 1883
- Macrochlamys japvoensis Godwin-Austen, 1910
- Macrochlamys kandiensis Godwin-Austen, 1883
- Macrochlamys kashmirensis (G.Nevill, 1878)
- Macrochlamys kasnakowi (Westerlund, 1898)
- Macrochlamys kelantanensis Möllendorff, 1902
- Macrochlamys kempi Godwin-Austen, 1883
- Macrochlamys koliaensis Godwin-Austen, 1883
- Macrochlamys kuluensis W.T.Blanford, 1904
- Macrochlamys kurtzi Godwin-Austen, 1910
- Macrochlamys lahupaensis Godwin-Austen, 1907
- Macrochlamys latus Godwin-Austen, 1888
- Macrochlamys lecythis (W.H.Benson, 1852)
- Macrochlamys leggeae Sajan, Tripathy, Chandra & Sivakumar, 2019
- Macrochlamys lemma Pholyotha & Panha, 2018
- Macrochlamys lemna Pholyotha & Panha, 2018
- Macrochlamys lhotaensis Godwin-Austen, 1883
- Macrochlamys lixa (W.T.Blanford, 1866)
- Macrochlamys loana (Gredler, 1882)
- Macrochlamys longicauda Godwin-Austen, 1883
- Macrochlamys lubrica (W.H.Benson, 1852)
- Macrochlamys lubricata Godwin-Austen, 1910
- Macrochlamys lumsdeni (Godwin-Austen, 1910)
- Macrochlamys luyorensis Godwin-Austen, 1918
- Macrochlamys luzonica (Möllendorff, 1894)
- Macrochlamys mahadeoensis Godwin-Austen, 1907
- Macrochlamys malaccana (L.Pfeiffer, 1855)
- Macrochlamys martini Schepman, 1893
- Macrochlamys masoni Godwin-Austen, 1910
- Macrochlamys mentaweiense Degner, 1928
- Macrochlamys minuta E.von Martens, 1891
- Macrochlamys molecula (W.H.Benson, 1859)
- Macrochlamys moupiniana (Deshayes, 1870)
- Macrochlamys munipurensis Godwin-Austen, 1907
- Macrochlamys murdochi Godwin-Austen, 1918
- Macrochlamys neherensis (W.H.Benson, 1864)
- Macrochlamys nemothaensis Godwin-Austen, 1910
- Macrochlamys nengloensis Godwin-Austen, 1883
- Macrochlamys nepos (L.Pfeiffer, 1855)
- Macrochlamys nitidissima Möllendorff, 1883
- Macrochlamys notha W.T.Blanford, 1904
- Macrochlamys novoguineensis Schepman, 1919
- Macrochlamys noxia W.T.Blanford, 1904
- Macrochlamys nuda (Reeve, 1852)
- Macrochlamys obiana Möllendorff, 1902
- Macrochlamys opiparus Godwin-Austen, 1883
- Macrochlamys originaria Godwin-Austen, 1883
- Macrochlamys pacata Godwin-Austen, 1883
- Macrochlamys papuana E.A.Smith, 1896
- Macrochlamys papuensis Preston, 1902
- Macrochlamys par Schmacker & Boettger, 1891
- Macrochlamys patane (W.H.Benson, 1859)
- Macrochlamys patens W.T.Blanford, 1904
- Macrochlamys paurhiensis (Godwin-Austen, 1899)
- Macrochlamys pedina (W.H.Benson, 1865)
- Macrochlamys pedisequae Heude, 1886
- Macrochlamys perfragilis Godwin-Austen, 1907
- Macrochlamys perfucata (W.H.Benson, 1853)
- Macrochlamys perincospicua Godwin-Austen, 1888
- Macrochlamys peringundensis R.H.Beddome, 1891
- Macrochlamys perplana Godwin-Austen, 1883
- Macrochlamys perrotteti (L.Pfeiffer, 1851)
- Macrochlamys petasus (W.H.Benson, 1859)
- Macrochlamys petrosa (T.Hutton, 1834)
- Macrochlamys planata (Heude, 1882)
- Macrochlamys planiuscula (W.H.Benson, 1838)
- Macrochlamys planorbiformis Bollinger, 1918
- Macrochlamys planula (Heude, 1882)
- Macrochlamys plicifera (W.T.Blanford, 1881)
- Macrochlamys politula Godwin-Austen, 1883
- Macrochlamys prava W.T.Blanford, 1905
- Macrochlamys proba Godwin-Austen, 1910
- Macrochlamys propinqua Godwin-Austen, 1910
- Macrochlamys pseudaulopis Godwin-Austen, 1888
- Macrochlamys pseudochoinix W.T.Blanford, 1904
- Macrochlamys pseudosuccinea E.A.Smith, 1894
- Macrochlamys pseustes E.A.Smith, 1893
- Macrochlamys psittacinus Godwin-Austen, 1918
- Macrochlamys psyche Vermeulen, Luu, Theary & Anker, 2019
- Macrochlamys quadrasi (Möllendorff, 1896)
- Macrochlamys quesita Godwin-Austen, 1910
- Macrochlamys rabani Godwin-Austen, 1910
- Macrochlamys radia Godwin-Austen, 1910
- Macrochlamys rakaensis Godwin-Austen, 1907
- Macrochlamys ramburianus (Mabille & Le Mesle, 1866)
- Macrochlamys razamiensis Godwin-Austen, 1907
- Macrochlamys rejecta (L.Pfeiffer, 1859)
- Macrochlamys rejectella (Mabille, 1889)
- Macrochlamys renitens (Morelet, 1861)
- Macrochlamys resinacea (Heude, 1885)
- Macrochlamys richilaensis Godwin-Austen, 1907
- Macrochlamys roberti Godwin-Austen, 1883
- Macrochlamys robsoni Rensch, 1932
- Macrochlamys rorida (W.H.Benson, 1859)
- Macrochlamys rotungensis Godwin-Austen, 1918
- Macrochlamys rubellocincta (W.T.Blanford, 1870)
- Macrochlamys russeola (Morelet, 1851)
- Macrochlamys rusticula Godwin-Austen, 1883
- Macrochlamys rutila W.T.Blanford, 1905
- Macrochlamys sacrata Godwin-Austen, 1910
- Macrochlamys salmonea (Ancey, 1882)
- Macrochlamys saltus Godwin-Austen, 1910
- Macrochlamys salwinensis Godwin-Austen, 1907
- Macrochlamys sanchezi Quadras & Möllendorff, 1894
- Macrochlamys sanctijohni (Godwin-Austen, 1891)
- Macrochlamys sata Godwin-Austen, 1883
- Macrochlamys sathilaensis Godwin-Austen, 1907
- Macrochlamys scyphus Godwin-Austen, 1910
- Macrochlamys segnis Pilsbry, 1934
- Macrochlamys sequax (W.H.Benson, 1859)
- Macrochlamys sequius Godwin-Austen, 1907
- Macrochlamys shengorensis Godwin-Austen, 1883
- Macrochlamys shimangensis Godwin-Austen, 1916
- Macrochlamys shisha (Godwin-Austen, 1875)
- Macrochlamys silcuriensis Godwin-Austen, 1914
- Macrochlamys silvatica Godwin-Austen, 1910
- Macrochlamys simplex Heude, 1890
- Macrochlamys sinensis (Heude, 1882)
- Macrochlamys sinica E.von Martens, 1875
- Macrochlamys sphaerica Godwin-Austen, 1910
- Macrochlamys spiralifer Vermeulen, 1996
- Macrochlamys spiriplana (Gredler, 1882)
- Macrochlamys spreta W.T.Blanford, 1904
- Macrochlamys stenogyra Möllendorff, 1901
- Macrochlamys stephoides Stoliczka, 1873
- Macrochlamys stephus (W.H.Benson, 1861)
- Macrochlamys strephoides Stoliczka, 1873
- Macrochlamys striata Godwin-Austen, 1910
- Macrochlamys striaticostata Godwin-Austen, 1910
- Macrochlamys stumpfii O.Boettger, 1889
- Macrochlamys subangulata Godwin-Austen, 1910
- Macrochlamys subcornea (L.Pfeiffer, 1861)
- Macrochlamys subcorpulenta Vermeulen & Liew, 2022
- Macrochlamys subjecta (W.H.Benson, 1852)
- Macrochlamys subpetasus Godwin-Austen, 1888
- Macrochlamys sufflava Godwin-Austen, 1910
- Macrochlamys superflua W.T.Blanford, 1904
- Macrochlamys superlita (Morelet, 1862)
- Macrochlamys sylhetensis Godwin-Austen, 1910
- Macrochlamys tanirensis Godwin-Austen, 1883
- Macrochlamys tanymentula Pholyotha & Panha, 2018
- Macrochlamys tecta (Souleyet, 1852)
- Macrochlamys tenuiarata Vermeulen & Liew, 2022
- Macrochlamys tenuicula H.Adams, 1868
- Macrochlamys tenuigranosa Dautzenberg, 1894
- Macrochlamys terminalis Godwin-Austen, 1910
- Macrochlamys terminus Godwin-Austen, 1899
- Macrochlamys tersa (Issel, 1874)
- Macrochlamys thaitieni Thach, 2021
- Macrochlamys theobaldi (Godwin-Austen, 1888)
- Macrochlamys tratanensis (Jousseaume, 1894)
- Macrochlamys trilobata Vermeulen & Liew, 2022
- Macrochlamys trusmadiensis Vermeulen & Liew, 2022
- Macrochlamys tugurium (W.H.Benson, 1852)
- Macrochlamys turanica E.von Martens, 1874
- Macrochlamys udus Godwin-Austen, 1899
- Macrochlamys umbraticola Godwin-Austen, 1883
- Macrochlamys umbrina (Reeve, 1854)
- Macrochlamys vallicola (Reeve, 1854)
- Macrochlamys vertex Godwin-Austen, 1910
- Macrochlamys vesica Godwin-Austen, 1910
- Macrochlamys vesicula (W.H.Benson, 1838)
- Macrochlamys vitrinoides (Deshayes, 1831)
- Macrochlamys woodiana (Reeve, 1852)
- Macrochlamys woodmasoni Godwin-Austen, 1888
- Macrochlamys zemoensis Godwin-Austen, 1907
- Macrochlamys zero (Mabille, 1887)
- 拟小囊螺属 Microcystina Mörch, 1872
- Oxytesta Zilch, 1956
- 罗氏螺属 Rahula Godwin-Austen, 1907
- 萨氏螺属 Sarika Godwin-Austen, 1907
- 赛氏螺属 Sesara Albers, 1860
- Syama W. T. Blanford & Godwin-Austen, 1908[19]
- 巨楯蛞蝓属 Tanychlamys Benson, 1834[4]
- Vitrinula Gray, 1857[4][20]
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小黄壳蛞蝓亚科 Ostracolethinae Simroth, 1901
- Cryptosemelus Collinge, 1902
- Laocaia Kuzminykh, 1999[4]
- Microparmarion Simroth, 1893[4]
- Microparmarion andamanica Collinge, 1901
- Microparmarion austeni Simroth, 1893
- Microparmarion basifixus Vermeulen & Liew, 2022
- Microparmarion boettgeri Simroth, 1898
- Microparmarion bruneopallescens Collinge, 1901
- Microparmarion convolutus Vermeulen & Liew, 2022
- Microparmarion exquadratus Schilthuizen, Clavera, Khoo, Bondar, Elder, Bouma, Eddington, Reisinger, Cosentino, Rossato & Delledonne, 2019
- Microparmarion fruhstorferi Simroth, 1898
- Microparmarion litteratus Schepman, 1896
- Microparmarion malayanus (Collinge, 1903)
- Microparmarion moellendorffi Collinge, 1901
- Microparmarion pollonerai Collinge & Godwin-Austen, 1895
- Microparmarion sallehi H.Wu, Ezzwan & Hamdani, 2023[22]
- Microparmarion simrothi Collinge & Godwin-Austen, 1895
- Microparmarion strubelli Simroth, 1893
- 米农螺属 Minyongia Godwin-Austen, 1916
- Minyongia kempi Godwin-Austen, 1916
- Myotesta Collinge, 1901[4]
- Ostracolethe Simroth, 1901 - 为Ostracolethinae亚科的模式属 [4]
- 鼈甲蛞蝓屬 Parmarion P. Fischer, 1855，盾勇螺属[3][4]
- 菲律宾蛞蝓属 Philippinella Möllendorff, 1899
- Philippinella carinata (Möllendorff, 1894)
- Philippinella moellendorffi Collinge, 1899
- Philippinella quadrasi (Möllendorff, 1894)
親緣支序圖
以下的系统发生学分支图顯示了根據Hausdorf (2000)所描述蛞蝓下目各科之間的親緣關係[6]：
| 蛞蝓下目 Limacoidei |
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參考文獻
- ^ 1.0 1.1 Godwin-Austen H. H. (April 1888). Land and freshwater Mollusca of India. including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean, supplementary to Messrs. Theobald and Hanley's Conchologia Indica. Taylor & Francis, London, volume 1, part 6: 207-257, page 253.
- ^ 中国科学院动物研究所. 真锥恰里螺. 《中国动物物种编目数据库》. 中国科学院微生物研究所. [2009-04-28]. （原始内容存档于2016-03-05）.
- ^ 3.0 3.1 3.2 Ariophantidae 絲鼈甲蝸牛科. 台灣生物多樣性資訊入口網. [2020-03-02]. （原始内容存档于2020-08-13） （中文（繁體））.
- ^ 4.00 4.01 4.02 4.03 4.04 4.05 4.06 4.07 4.08 4.09 4.10 Bouchet, P.; Rocroi, J.-P.; Frýda, J.; Hausdorf, B.; Ponder, W.; Valdés, Á. & Warén, A. Classification and nomenclator of gastropod families. Malacologia (Hackenheim, Germany: ConchBooks). 2005, 47 (1–2): 1–397. ISBN 3-925919-72-4. OCLC 61351079. OL 42989175M. S2CID 83847814.
- ^ Bouchet, P.; Rocroi, J.-P.; Hausdorf, B.; Kaim, A.; Kano, Y.; Nützel, A.; Parkhaev, P.; Schrödl, M.; Strong, E.E. Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families. Malacologia. 2017, 61 (1–2): 1–526. OCLC 7296100651. S2CID 91051256. doi:10.4002/040.061.0201.
- ^ 6.0 6.1 Hausdorf, B. Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal. Journal of Biogeography. 2000, 27 (2): 379–390. doi:10.1046/j.1365-2699.2000.00403.x （英语）.
- ^ Gude G. P. L. K. & Woodward B. B. (1921). "On Helicella, Férussac". Proceedings of the Malacological Society of London 14(5-6): 174-190. page 186.
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延伸閱讀
- Schileyko A. A. (2003). "Treatise on recent terrestrial pulmonate mollusks. 10. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae". Ruthenica, Supplement 2. 1309-1466.
